Roger (Mac) Strand


Contact Information:

2121 Weston Hall
Phone (906) 227-2365


  • Michigan State University, Ph.D. 1996
  • University of Minnesota, M.S. 1990
  • Gustavus Adolphus College, B.A. 1985

Research Interests:

Ecology of the Lake Superior Wave Zone
The coastal waters of Lake Superior are home to a community of organisms uniquely adapted to face the challenges of life in the often powerful freshwater surf.  My research in the Lake Superior wave zone is focused on investigating species interactions, trophic structuring, and energy flow in wave zone and tributary communities.

My colleagues from the United States Geological Survey in Wisconsin and I are investigating the role of groundwater-surface water exchange dynamics in structuring northwoods stream communities and governing productivity through controls on nutrient delivery to and from streambed sediments. 


  • BI 465 Aquatic Insect Ecology
  • BI 424 General Entomology
  • BI 305 Ecology of the Northern Forest
  • BI 410 Ecology of the Great Lakes
  • BI 495 Stream Ecology
  • BI 324 Invertebrate Zoology

Selected Publications:

  • Skelton, J. and M. Strand. 2013. Trophic ecology of a freshwater sponge (Spongilla lacustris) revealed by stable isotope analysis.  Hydrobiologia 709: 227-235. PDF
  • Hunt, R.J., M. Strand, and J.F. Walker.  2006.  Measuring groundwater-surface water interaction and its effect on wetland stream benthic productivity, Trout Lake watershed, northern Wisconsin, USA.  Journal of Hydrology 320: 370-384. PDF
  • Strand, M.  2005.  Trophic ecology of the Lake Superior wave zone: a stable isotope approach.  Hydrobiologia 544: 271-278. PDF
  • Strand, R.M. and R.W. Merritt.  1999.  Impacts of livestock grazing activities on stream insect communities and the riverine environment.  The American Entomologist 45: 13-29.
  • Strand, M., D.A. Herms, M.P. Ayres, M.E. Kubiske, M.G. Kaufman, E.D. Walker, K.S. Pregitzer, and R.W. Merritt.  1999.  Effects of atmospheric CO2, light availability, and tree species on the quality of leaf detritus as a resource for treehole mosquitoes.  Oikos 84: 277-283.
Roger (Mac) Strand